Preface |
|
xiii | |
Author Biography |
|
xvii | |
Abbreviations |
|
xix | |
|
Chapter 1 Introduction to Phylogenetics |
|
|
1 | (62) |
|
1.1 Logical and Conceptual Aspects of Phylogenetic Analysis |
|
|
2 | (6) |
|
|
3 | (3) |
|
|
6 | (1) |
|
|
7 | (1) |
|
|
8 | (3) |
|
|
9 | (2) |
|
1.3 Parsimony, Synapomorphies, and Rooting |
|
|
11 | (4) |
|
1.4 Maximum Likelihood and Assumptions Implicit in Parsimony |
|
|
15 | (3) |
|
1.5 Distances, Phenetics, and Information Content |
|
|
18 | (9) |
|
1.5.1 Distances and Their Properties |
|
|
19 | (3) |
|
1.5.2 Phenetics vs. Cladistics |
|
|
22 | (1) |
|
1.5.3 Retrieving Distances |
|
|
23 | (2) |
|
1.5.4 Transmitting Character Information |
|
|
25 | (2) |
|
1.6 Pattern Cladistics and Three-Taxon Statements |
|
|
27 | (4) |
|
1.6.1 Three-Taxon Statements |
|
|
28 | (3) |
|
1.7 Phylogeny as an Assumption; Limits of Phylogeny |
|
|
31 | (3) |
|
1.8 Optimality Criterion and Foundations of Phylogenetic Analysis |
|
|
34 | (2) |
|
1.9 On the Need for Computer Programs |
|
|
36 | (1) |
|
1.10 Implementation in TNT: Tree Analysis Using New Technology |
|
|
37 | (3) |
|
1.11 Input, Commands, Format of Data Matrices |
|
|
40 | (16) |
|
1.11.1 Commands and Truncation |
|
|
40 | (2) |
|
|
42 | (1) |
|
|
43 | (1) |
|
1.11.4 Lists and Ranges; Numbering of Elements |
|
|
43 | (2) |
|
1.11.5 Reading Basic Data |
|
|
45 | (1) |
|
1.11.6 Different Data Formats and Sizes |
|
|
46 | (2) |
|
1.11.7 Wiping the Dataset from Memory |
|
|
48 | (1) |
|
1.11.8 Multiple Blocks and Data Formats |
|
|
48 | (2) |
|
1.11.9 Combining Existing Datasets |
|
|
50 | (1) |
|
1.11.10 Datasets in Multiple Files |
|
|
50 | (2) |
|
1.11.11 Redefining Data Blocks |
|
|
52 | (1) |
|
1.11.12 Other Data Formats |
|
|
52 | (1) |
|
1.11.13 Changing the Data |
|
|
53 | (2) |
|
1.11.14 Saving Edited Dataset in Different Formats |
|
|
55 | (1) |
|
|
56 | (4) |
|
|
56 | (1) |
|
1.12.2 GUI Screens (Windows Only) |
|
|
56 | (1) |
|
1.12.3 Outputting Numbers or Names |
|
|
57 | (1) |
|
|
57 | (1) |
|
|
58 | (1) |
|
|
58 | (1) |
|
|
59 | (1) |
|
1.12.8 Progress Reports, Warnings |
|
|
59 | (1) |
|
|
59 | (1) |
|
1.12.10 Saving Trees to Files |
|
|
60 | (1) |
|
1.13 Outline of the Remaining Chapters |
|
|
60 | (3) |
|
Chapter 2 Characters, Homology, and Datasets |
|
|
63 | (56) |
|
2.1 The Great Chain of Characters |
|
|
63 | (1) |
|
|
64 | (4) |
|
2.2.1 Two Main Meanings of Homology |
|
|
65 | (2) |
|
|
67 | (1) |
|
2.3 Criteria for Homology |
|
|
68 | (4) |
|
2.4 Homology by Special Knowledge? |
|
|
72 | (2) |
|
2.5 No Special Knowledge of Homology Is Possible or Necessary |
|
|
74 | (3) |
|
2.6 Life Stages, Comparability, Ontogeny |
|
|
77 | (3) |
|
2.7 Gathering Morphological Data |
|
|
80 | (5) |
|
2.8 Character Independence |
|
|
85 | (3) |
|
|
88 | (2) |
|
2.10 Character Coding and Character Types |
|
|
90 | (7) |
|
2.10.1 Discrete Characters |
|
|
90 | (7) |
|
2.11 Transformation Series Analysis |
|
|
97 | (1) |
|
2.12 Continuous and Landmark Data |
|
|
98 | (2) |
|
|
100 | (1) |
|
|
101 | (18) |
|
2.14.1 Basic Character Settings: ccode Command |
|
|
101 | (2) |
|
2.14.2 Step-Matrix Characters (and Ancestral States) |
|
|
103 | (6) |
|
2.14.3 Deactivating Blocks of Data |
|
|
109 | (1) |
|
|
109 | (2) |
|
2.14.5 Taxon Settings and Taxonomic Information |
|
|
111 | (4) |
|
2.14.6 Comparing and Merging Datasets |
|
|
115 | (4) |
|
Chapter 3 Character Optimization: Evaluation of Trees and Inference of Ancestral States |
|
|
119 | (42) |
|
3.1 Finding Optimal Ancestral Reconstructions |
|
|
119 | (2) |
|
3.2 Generalized Optimization: Simple Cases |
|
|
121 | (3) |
|
3.3 Optimization for Nonadditive Characters: Fitch's (1971) Method |
|
|
124 | (3) |
|
3.4 Optimization for Additive Characters: Farris's (1970) Method |
|
|
127 | (5) |
|
3.5 Step-Matrix Optimization |
|
|
132 | (3) |
|
3.6 Other Types of Optimization |
|
|
135 | (1) |
|
3.7 Ambiguity, Polymorphisms, Missing Entries |
|
|
135 | (3) |
|
|
137 | (1) |
|
|
138 | (1) |
|
3.8 Mapping, Synapomorphies, and Reconstructed Ancestors |
|
|
138 | (3) |
|
3.8.1 Reconstructed Ancestors |
|
|
140 | (1) |
|
|
141 | (1) |
|
|
141 | (2) |
|
3.10 Polytomies, Multiple MPTs, and Consensus |
|
|
143 | (4) |
|
3.10.1 Length of Polytomies and Their Resolutions |
|
|
143 | (1) |
|
3.10.2 Polytomies as "Soft" |
|
|
144 | (1) |
|
3.10.3 Informative Characters |
|
|
145 | (1) |
|
3.10.4 Mapping Multiple Trees |
|
|
145 | (2) |
|
|
147 | (4) |
|
3.12 Readability of Ancestors |
|
|
151 | (2) |
|
3.13 Implementation in TNT |
|
|
153 | (8) |
|
3.13.1 Options for Scoring Trees |
|
|
153 | (2) |
|
3.13.2 Diagnosis and Mapping |
|
|
155 | (1) |
|
3.13.3 Diagrams for Publication |
|
|
156 | (1) |
|
3.13.4 Reconstructions and Specific Changes |
|
|
157 | (1) |
|
3.13.5 Selecting and Preparing the Trees to Be Optimized |
|
|
158 | (3) |
|
Chapter 4 Models and Assumptions in Morphology |
|
|
161 | (46) |
|
4.1 Maximum Likelihood (ML) |
|
|
162 | (1) |
|
4.2 Assumptions of Models of Molecular Evolution |
|
|
163 | (4) |
|
4.3 Likelihood Calculation |
|
|
167 | (4) |
|
|
167 | (2) |
|
|
169 | (1) |
|
|
170 | (1) |
|
4.4 Among Site Rate Variation |
|
|
171 | (1) |
|
4.5 Linked and Unlinked Partitions |
|
|
172 | (1) |
|
|
173 | (2) |
|
4.7 Some Difficulties with Bayesian Phylogenetics |
|
|
175 | (5) |
|
4.7.1 No Optimality Criterion |
|
|
176 | (1) |
|
|
176 | (1) |
|
4.7.3 Summarizing Results |
|
|
176 | (4) |
|
4.7.4 Sample Size and Frequency |
|
|
180 | (1) |
|
|
180 | (1) |
|
|
180 | (2) |
|
4.9 Adapting Models for Molecular evolution to Morphology |
|
|
182 | (4) |
|
|
183 | (1) |
|
|
183 | (1) |
|
4.9.3 Assumptions of Mk/Mkv Models |
|
|
184 | (2) |
|
4.10 Parsimony, Models, and Consistency |
|
|
186 | (9) |
|
4.10.1 Low Rates of Change in the MDG Make Parsimony Consistent |
|
|
187 | (1) |
|
4.10.2 Inferring Trees by Fixing Branch Lengths and Using Best Individual Reconstruction Amounts to Parsimony |
|
|
187 | (1) |
|
4.10.3 For Data Generated with All Branches of the Same Length, Parsimony Produces Good Results |
|
|
188 | (3) |
|
4.10.4 If All Characters and Branches Can Have Different Lengths, MP Is Identical to ML |
|
|
191 | (2) |
|
4.10.5 Invariant Characters and a Large Number of States |
|
|
193 | (2) |
|
4.10.6 Missing Data and Likelihood |
|
|
195 | (1) |
|
4.11 Standard Poisson Models in Morphology |
|
|
195 | (7) |
|
|
200 | (2) |
|
|
202 | (1) |
|
|
203 | (4) |
|
Chapter 5 Tree Searches: Finding Most Parsimonious Trees |
|
|
207 | (48) |
|
|
208 | (3) |
|
5.2 Small Datasets: Exact Solutions |
|
|
211 | (1) |
|
5.3 Datasets of Medium Difficulty: Basic Methods |
|
|
212 | (9) |
|
|
212 | (2) |
|
|
214 | (2) |
|
|
216 | (1) |
|
5.3.4 Local Optima and Islands |
|
|
216 | (2) |
|
5.3.5 Escaping Local Optima |
|
|
218 | (1) |
|
5.3.6 Comparing Efficiency of Search Algorithms |
|
|
219 | (2) |
|
5.4 Datasets of Medium Difficulty: Multiple Starting Points in Depth |
|
|
221 | (6) |
|
5.4.1 Wagner Trees vs. Random Trees |
|
|
221 | (1) |
|
5.4.2 Saving Reduced Numbers of Trees per Replication |
|
|
221 | (1) |
|
5.4.3 Effect of Full Tree Buffer |
|
|
222 | (1) |
|
5.4.4 Convergence and Choice of Search Settings |
|
|
223 | (1) |
|
5.4.5 Collapsing of Zero-Length Branches and Search Efficiency |
|
|
224 | (2) |
|
5.4.6 Many Hits or Many Trees? |
|
|
226 | (1) |
|
5.5 Searches under Constraints; Backbone Topologies |
|
|
227 | (2) |
|
5.6 Difficult Datasets: Basic Ideas and Methods |
|
|
229 | (11) |
|
|
229 | (1) |
|
5.6.2 Sectorial Searches (55) |
|
|
230 | (1) |
|
5.6.2.1 Types of Sector Selection |
|
|
231 | (2) |
|
5.6.2.2 Analysis of Reduced Datasets |
|
|
233 | (1) |
|
|
233 | (2) |
|
|
235 | (1) |
|
|
236 | (1) |
|
|
236 | (2) |
|
|
238 | (1) |
|
|
238 | (2) |
|
|
240 | (1) |
|
5.7 Difficult Datasets: Combined Methods and Driven Searches |
|
|
240 | (3) |
|
5.7.1 Searching for a Stable Consensus |
|
|
241 | (1) |
|
5.7.2 Strengths and Weaknesses of the Different Algorithms |
|
|
241 | (1) |
|
5.7.2.1 Alternatives to the Algorithms Described |
|
|
242 | (1) |
|
5.7.3 Challenges Posed by Morphological Datasets |
|
|
243 | (1) |
|
5.8 Approximate Searches and Quick Consensus Estimation |
|
|
243 | (3) |
|
5.9 Implementation in TNT |
|
|
246 | (9) |
|
|
247 | (2) |
|
|
249 | (1) |
|
5.9.3 Heuristic Searches--Basic Algorithms |
|
|
249 | (1) |
|
5.9.4 Heuristic Searches--Special Algorithms |
|
|
250 | (5) |
References |
|
255 | (18) |
Index |
|
273 | |